He ["the male"] is trapped in a twilight zone halfway...

He ["the male"] is trapped in a twilight zone halfway between humans and apes, and is far worse off than apes, because he is, first of all, capable of a large array of negative feelings that the apes aren't - hate, jealousy, contempt, disgust, guilt, shame, disgrace, doubt - and, secondly, he is aware of what he is and isn't.